Present theories for predicting expected Monte Carlo errors in neutron transport calculations apply to estimates of flux-weighted integrals sampled directly by scoring individual collisions. To treat track-length estimators, the recent theory of Amster and Djomehri is generalized to allow the score distribution functions to depend on the coordinates of two successive collisions. It has long been known that the expected track length in a region of phase space equals the expected flux integrated over that region, but that the expected statistical error of the Monte Carlo estimate of the track length is different from that of the flux integral obtained by sampling the sum of the reciprocals of the total cross sections for all collisions in the region. These conclusions are shown to be implied by the generalized theory, which provides explicit equations for the expected values and errors of both types of estimators. Sampling expected contributions to the track-length estimator is also treated. Other general properties of the errors for both estimators are derived from the equations and physically interpreted. The actual values of these errors are then obtained and interpreted for a simple specific example.